01 / Morfologi

Ciri dan struktur

Associated with twigs and branches of Ulmus pumila L. (Ulmaceae). Sexual morph Undetermined. Asexual morph Conidiostromata 1000–1700 × 500–800 µm diam., semi-immersed in host tissue, solitary, erumpent, scattered, discoid, circular to ovoid, with multi-loculate, pycnidial, embedded in stromatic tissue, with ostiole. Ostioles 200–500 µm long, with an ostiolar neck. Conidiomata walls comprising a few layers of pseudoparenchymatous cells of textura angularis, with inner layer thin, pale brown, outer layer brown to dark brown. Conidiophores unbranched or occasionally branched at the base, formed from the inner layer of the pycnidial wall. Conidiogenous cells (6–)9–14 × 1.3–1.8(–2.8) μm (\(\bar{x} \) = 11 × 2 μm, n = 15), blastic, enteroblastic, flask-shaped, phialidic, hyaline, and smooth-walled. Conidia (7–)8–9.5 × 1.5–1.7(–1.9) µm (\(\bar{x} \) = 8.5 × 1.6 µm, n = 30), unicellular, hyaline, oblong to allantoid, smooth-walled. Culture characteristics: Conidia germinating on PDA within 12 h. Germ tubes produced from all sides. Colonies on PDA reaching 22–27 mm diam. after 2 weeks at room temperature, colonies irregular in shape, medium dense, flat to slightly raised, surface slightly rough, with small granular, to velvety, edge undulate with well-defined margin, from above pale orange to light brown at the margin, amber near the margin, with, zonate, pale yellowish to dandelion and dark amber at the centre; from below pale yellowish at the margin, with dark amber at the centre; not producing pigmentation in agar.

Sebagian uraian ditampilkan dalam bahasa sumber; terjemahannya belum tersedia atau belum lolos pemeriksaan.

Rujukan: [1]

Pengukuran mengikuti objek, spesimen atau kultur, dan metode yang disebutkan sumber. Baca uraian asli untuk konteks lengkap.

02 / Kehidupan

Habitat dan peran ekologis

Culture characteristics: Conidia germinating on PDA within 12 h. Germ tubes produced from all sides. Colonies on PDA reaching 22–27 mm diam. after 2 weeks at room temperature, colonies irregular in shape, medium dense, flat to slightly raised, surface slightly rough, with small granular, to velvety, edge undulate with well-defined margin, from above pale orange to light brown at the margin, amber near the margin, with, zonate, pale yellowish to dandelion and dark amber at the centre; from below pale yellowish at the margin, with dark amber at the centre; not producing pigmentation in agar. Material examined: RUSSIA, Rostov region, Shakhty City, street trees, on dead branch of Ulmus pumila L. (Ulmaceae), 27 May 2017, T.S. Bulgakov, T-1778 (MFLU 17-2080, holotype; PDD, isotype), ex-type living culture, MFLUCC 18-1227. Notes: Cytospora ulmicola (MFLU 17-2080) was found on a dead branch (with signs of necrosis) of Ulmus pumila in European Russia. Cytospora ulmicola resembles Cytospora species in having an ostiolar neck with multi-loculate conidiostromata and unicellular, oblong to allantoid conidia. Phylogenetic analysis based on a combined ITS, LSU and ACT sequence dataset indicates that C. ulmicola clusters with C. cotini Norph. et al. (MFLUCC 14-1050), C. ampulliformis Norph. et al. (MFLUCC 16-0583), C. gelida Norph. et al. (MFLUCC 16-0634), and C. ceratosperma (Tode) G.C.

Sebagian uraian ditampilkan dalam bahasa sumber; terjemahannya belum tersedia atau belum lolos pemeriksaan.

Rujukan: [1]

Habitat atau konteks temuan
kultur yang didokumentasikan
Substrat
Informasi belum tersedia pada sumber rilis ini.
Peran atau konteks
Belum dikategorikan pada rilis ini.
Temukan hubungan serupa
03 / Siklus hidup

Reproduksi

Associated with twigs and branches of Ulmus pumila L. (Ulmaceae). Sexual morph Undetermined. Asexual morph Conidiostromata 1000–1700 × 500–800 µm diam., semi-immersed in host tissue, solitary, erumpent, scattered, discoid, circular to ovoid, with multi-loculate, pycnidial, embedded in stromatic tissue, with ostiole. Ostioles 200–500 µm long, with an ostiolar neck. Conidiomata walls comprising a few layers of pseudoparenchymatous cells of textura angularis, with inner layer thin, pale brown, outer layer brown to dark brown. Conidiophores unbranched or occasionally branched at the base, formed from the inner layer of the pycnidial wall. Conidiogenous cells (6–)9–14 × 1.3–1.8(–2.8) μm (\(\bar{x} \) = 11 × 2 μm, n = 15), blastic, enteroblastic, flask-shaped, phialidic, hyaline, and smooth-walled.

Sebagian uraian ditampilkan dalam bahasa sumber; terjemahannya belum tersedia atau belum lolos pemeriksaan.

Rujukan: [1]

04 / Batas informasi

Catatan identitas dan cakupan

  • Nama diterima mengikuti Catalogue of Life Base2026-09-11; uraian mempertahankan konsep dan nama sumber aslinya.
  • Preserve original monograph treatment localities, hosts and specimen/substrate scope. Materials examined document specimens, not the full global range.
  • Impor dan terjemahan otomatis belum diperiksa editorial per profil. Struktur reproduktif yang disebutkan tidak membuktikan seluruh siklus hidup diketahui.
  • Pengukuran pada uraian mengikuti spesimen, kultur, dan metode sumber; angka tidak diperluas ke semua populasi.

Catatan distribusi dalam narasi mengikuti sumbernya. Dokumentasi spesimen atau foto mengikuti catatan sumber. Bukti wilayah Indonesia dan penilaian konservasi tidak disimpulkan dari gambar.

05 / Dokumentasi

Melihat dengan konteks

Cytospora ulmicola (MFLU 17-2080, holotype). a Herbarium label and specimens. b, d, e Appearance of conidiostromata on branches of Ulmus pumila. c, f Colonies on PDA (c = from above, f = from below). g Transverse sections through conidiostroma to show an arrangement of the locules. h Longitudinal section through conidiostroma. i Ostiolar neck. j Conidiostroma wall. k Conidiogenous cells. l Conidia. Scale bars b = 1000 µm, e = 100 µm, g, h = 200 µm, i = 50 µm, j = 20 µm, k, l = 10 µm

Jenis
Pelat ilmiah
Kredit
Penulis dan sumber artikel: Phookamsak, Rungtiwa; Hyde, Kevin D.; Jeewon, Rajesh; Bhat, D. Jayarama; Jones, E. B. Gareth; Maharachchikumbura, Sajeewa S. N.; Raspé, Olivier; Karunarathna, Samantha C.; Wanasinghe, Dhanushka N.; Hongsanan, Sinang; Doilom, Mingkwan; Tennakoon, Danushka S.; Machado, Alexandre R.; Firmino, André L.; Ghosh, Aniket; Karunarathna, Anuruddha; Mešić, Armin; Dutta, Arun Kumar; Thongbai, Benjarong; Devadatha, Bandarupalli; Norphanphoun, Chada; Senwanna, Chanokned; Wei, Deping; Pem, Dhandevi; Ackah, Frank Kwekucher; Wang, Gen-Nuo; Jiang, Hong-Bo; Madrid, Hugo; Lee, Hyang Burm; Goonasekara, Ishani D.; Manawasinghe, Ishara S.; Kušan, Ivana; Cano, Josep; Gené, Josepa; Li, Junfu; Das, Kanad; Acharya, Krishnendu; Raj, K. N. Anil; Latha, K. P. Deepna; Chethana, K. W. Thilini; He, Mao-Qiang; Dueñas, Margarita; Jadan, Margita; Martín, María P.; Samarakoon, Milan C.; Dayarathne, Monika C.; Raza, Mubashar; Park, Myung Soo; Telleria, M. Teresa; Chaiwan, Napalai; Matočec, Neven; de Silva, Nimali I.; Pereira, Olinto L.; Singh, Paras Nath; Manimohan, Patinjareveettil; Uniyal, Priyanka; Shang, Qiu-Ju; Bhatt, Rajendra P.; Perera, Rekhani H.; Alvarenga, Renato Lúcio Mendes; Nogal-Prata, Sandra; Singh, Sanjay K.; Vadthanarat, Santhiti; Oh, Seung-Yoon; Huang, Shi-Ke; Rana, Shiwali; Konta, Sirinapa; Paloi, Soumitra; Jayasiri, Subashini C.; Jeon, Sun Jeong; Mehmood, Tahir; Gibertoni, Tatiana Baptista; Nguyen, Thuong T. T.; Singh, Upendra; Thiyagaraja, Vinodhini; Sarma, V. Venkateswara; Dong, Wei; Yu, Xian-Dong; Lu, Yong-Zhong; Lim, Young Woon; Chen, Yun; Tkalčec, Zdenko; Zhang, Zhi-Feng; Luo, Zong-Long; Daranagama, Dinushani A.; Thambugala, Kasun M.; Tibpromma, Saowaluck; Camporesi, Erio; Bulgakov, Timur S.; Dissanayake, Asha J.; Senanayake, Indunil C.; Dai, Dong Qin; Tang, Li-Zhou; Khan, Sehroon; Zhang, Huang; Promputtha, Itthayakorn; Cai, Lei; Chomnunti, Putarak; Zhao, Rui-Lin; Lumyong, Saisamorn; Boonmee, Saranyaphat; Wen, Ting-Chi; Mortimer, Peter E.; Xu, Jianchu
Lisensi
CC BY 4.0
Sumber media
Buka entri sumber

Gambar menunjukkan objek menurut identifikasi sumber, bukan verifikasi independen spesimen oleh Fungi. Tidak ada skala baru yang ditambahkan.

06 / Rujukan

Sumber profil

  1. [1] Fungal diversity notes 929–1035: taxonomic and phylogenetic contributions on genera and species of fungi — Cytospora ulmicolaBagian sumber: publisher:10.1007/s13225-019-00421-w:paragraph:849
    Tanggal penghimpunan sumber: 2026-10-12
    Teks sumber: CC BY 4.0 · Phookamsak, Rungtiwa; Hyde, Kevin D.; Jeewon, Rajesh; Bhat, D. Jayarama; Jones, E. B. Gareth; Maharachchikumbura, Sajeewa S. N.; Raspé, Olivier; Karunarathna, Samantha C.; Wanasinghe, Dhanushka N.; Hongsanan, Sinang; Doilom, Mingkwan; Tennakoon, Danushka S.; Machado, Alexandre R.; Firmino, André L.; Ghosh, Aniket; Karunarathna, Anuruddha; Mešić, Armin; Dutta, Arun Kumar; Thongbai, Benjarong; Devadatha, Bandarupalli; Norphanphoun, Chada; Senwanna, Chanokned; Wei, Deping; Pem, Dhandevi; Ackah, Frank Kwekucher; Wang, Gen-Nuo; Jiang, Hong-Bo; Madrid, Hugo; Lee, Hyang Burm; Goonasekara, Ishani D.; Manawasinghe, Ishara S.; Kušan, Ivana; Cano, Josep; Gené, Josepa; Li, Junfu; Das, Kanad; Acharya, Krishnendu; Raj, K. N. Anil; Latha, K. P. Deepna; Chethana, K. W. Thilini; He, Mao-Qiang; Dueñas, Margarita; Jadan, Margita; Martín, María P.; Samarakoon, Milan C.; Dayarathne, Monika C.; Raza, Mubashar; Park, Myung Soo; Telleria, M. Teresa; Chaiwan, Napalai; Matočec, Neven; de Silva, Nimali I.; Pereira, Olinto L.; Singh, Paras Nath; Manimohan, Patinjareveettil; Uniyal, Priyanka; Shang, Qiu-Ju; Bhatt, Rajendra P.; Perera, Rekhani H.; Alvarenga, Renato Lúcio Mendes; Nogal-Prata, Sandra; Singh, Sanjay K.; Vadthanarat, Santhiti; Oh, Seung-Yoon; Huang, Shi-Ke; Rana, Shiwali; Konta, Sirinapa; Paloi, Soumitra; Jayasiri, Subashini C.; Jeon, Sun Jeong; Mehmood, Tahir; Gibertoni, Tatiana Baptista; Nguyen, Thuong T. T.; Singh, Upendra; Thiyagaraja, Vinodhini; Sarma, V. Venkateswara; Dong, Wei; Yu, Xian-Dong; Lu, Yong-Zhong; Lim, Young Woon; Chen, Yun; Tkalčec, Zdenko; Zhang, Zhi-Feng; Luo, Zong-Long; Daranagama, Dinushani A.; Thambugala, Kasun M.; Tibpromma, Saowaluck; Camporesi, Erio; Bulgakov, Timur S.; Dissanayake, Asha J.; Senanayake, Indunil C.; Dai, Dong Qin; Tang, Li-Zhou; Khan, Sehroon; Zhang, Huang; Promputtha, Itthayakorn; Cai, Lei; Chomnunti, Putarak; Zhao, Rui-Lin; Lumyong, Saisamorn; Boonmee, Saranyaphat; Wen, Ting-Chi; Mortimer, Peter E.; Xu, Jianchu
  2. [2] Catalogue of Life Base2026-09-11 — Cytospora ulmicolaBagian sumber: ColDP NameUsage.tsv ID 33SK8; accepted species; source 2073
    Tanggal penghimpunan sumber: 2026-10-12
  3. [3] Dokumentasi visual — Cytospora ulmicolaBagian sumber: Individual media record; identification follows the source, not independent determination
    Tanggal penghimpunan sumber: 2026-10-12
    Media sumber: CC BY 4.0 · Penulis dan sumber artikel: Phookamsak, Rungtiwa; Hyde, Kevin D.; Jeewon, Rajesh; Bhat, D. Jayarama; Jones, E. B. Gareth; Maharachchikumbura, Sajeewa S. N.; Raspé, Olivier; Karunarathna, Samantha C.; Wanasinghe, Dhanushka N.; Hongsanan, Sinang; Doilom, Mingkwan; Tennakoon, Danushka S.; Machado, Alexandre R.; Firmino, André L.; Ghosh, Aniket; Karunarathna, Anuruddha; Mešić, Armin; Dutta, Arun Kumar; Thongbai, Benjarong; Devadatha, Bandarupalli; Norphanphoun, Chada; Senwanna, Chanokned; Wei, Deping; Pem, Dhandevi; Ackah, Frank Kwekucher; Wang, Gen-Nuo; Jiang, Hong-Bo; Madrid, Hugo; Lee, Hyang Burm; Goonasekara, Ishani D.; Manawasinghe, Ishara S.; Kušan, Ivana; Cano, Josep; Gené, Josepa; Li, Junfu; Das, Kanad; Acharya, Krishnendu; Raj, K. N. Anil; Latha, K. P. Deepna; Chethana, K. W. Thilini; He, Mao-Qiang; Dueñas, Margarita; Jadan, Margita; Martín, María P.; Samarakoon, Milan C.; Dayarathne, Monika C.; Raza, Mubashar; Park, Myung Soo; Telleria, M. Teresa; Chaiwan, Napalai; Matočec, Neven; de Silva, Nimali I.; Pereira, Olinto L.; Singh, Paras Nath; Manimohan, Patinjareveettil; Uniyal, Priyanka; Shang, Qiu-Ju; Bhatt, Rajendra P.; Perera, Rekhani H.; Alvarenga, Renato Lúcio Mendes; Nogal-Prata, Sandra; Singh, Sanjay K.; Vadthanarat, Santhiti; Oh, Seung-Yoon; Huang, Shi-Ke; Rana, Shiwali; Konta, Sirinapa; Paloi, Soumitra; Jayasiri, Subashini C.; Jeon, Sun Jeong; Mehmood, Tahir; Gibertoni, Tatiana Baptista; Nguyen, Thuong T. T.; Singh, Upendra; Thiyagaraja, Vinodhini; Sarma, V. Venkateswara; Dong, Wei; Yu, Xian-Dong; Lu, Yong-Zhong; Lim, Young Woon; Chen, Yun; Tkalčec, Zdenko; Zhang, Zhi-Feng; Luo, Zong-Long; Daranagama, Dinushani A.; Thambugala, Kasun M.; Tibpromma, Saowaluck; Camporesi, Erio; Bulgakov, Timur S.; Dissanayake, Asha J.; Senanayake, Indunil C.; Dai, Dong Qin; Tang, Li-Zhou; Khan, Sehroon; Zhang, Huang; Promputtha, Itthayakorn; Cai, Lei; Chomnunti, Putarak; Zhao, Rui-Lin; Lumyong, Saisamorn; Boonmee, Saranyaphat; Wen, Ting-Chi; Mortimer, Peter E.; Xu, Jianchu

Adaptasi teks: CC BY 4.0. Phookamsak, Rungtiwa; Hyde, Kevin D.; Jeewon, Rajesh; Bhat, D. Jayarama; Jones, E. B. Gareth; Maharachchikumbura, Sajeewa S. N.; Raspé, Olivier; Karunarathna, Samantha C.; Wanasinghe, Dhanushka N.; Hongsanan, Sinang; Doilom, Mingkwan; Tennakoon, Danushka S.; Machado, Alexandre R.; Firmino, André L.; Ghosh, Aniket; Karunarathna, Anuruddha; Mešić, Armin; Dutta, Arun Kumar; Thongbai, Benjarong; Devadatha, Bandarupalli; Norphanphoun, Chada; Senwanna, Chanokned; Wei, Deping; Pem, Dhandevi; Ackah, Frank Kwekucher; Wang, Gen-Nuo; Jiang, Hong-Bo; Madrid, Hugo; Lee, Hyang Burm; Goonasekara, Ishani D.; Manawasinghe, Ishara S.; Kušan, Ivana; Cano, Josep; Gené, Josepa; Li, Junfu; Das, Kanad; Acharya, Krishnendu; Raj, K. N. Anil; Latha, K. P. Deepna; Chethana, K. W. Thilini; He, Mao-Qiang; Dueñas, Margarita; Jadan, Margita; Martín, María P.; Samarakoon, Milan C.; Dayarathne, Monika C.; Raza, Mubashar; Park, Myung Soo; Telleria, M. Teresa; Chaiwan, Napalai; Matočec, Neven; de Silva, Nimali I.; Pereira, Olinto L.; Singh, Paras Nath; Manimohan, Patinjareveettil; Uniyal, Priyanka; Shang, Qiu-Ju; Bhatt, Rajendra P.; Perera, Rekhani H.; Alvarenga, Renato Lúcio Mendes; Nogal-Prata, Sandra; Singh, Sanjay K.; Vadthanarat, Santhiti; Oh, Seung-Yoon; Huang, Shi-Ke; Rana, Shiwali; Konta, Sirinapa; Paloi, Soumitra; Jayasiri, Subashini C.; Jeon, Sun Jeong; Mehmood, Tahir; Gibertoni, Tatiana Baptista; Nguyen, Thuong T. T.; Singh, Upendra; Thiyagaraja, Vinodhini; Sarma, V. Venkateswara; Dong, Wei; Yu, Xian-Dong; Lu, Yong-Zhong; Lim, Young Woon; Chen, Yun; Tkalčec, Zdenko; Zhang, Zhi-Feng; Luo, Zong-Long; Daranagama, Dinushani A.; Thambugala, Kasun M.; Tibpromma, Saowaluck; Camporesi, Erio; Bulgakov, Timur S.; Dissanayake, Asha J.; Senanayake, Indunil C.; Dai, Dong Qin; Tang, Li-Zhou; Khan, Sehroon; Zhang, Huang; Promputtha, Itthayakorn; Cai, Lei; Chomnunti, Putarak; Zhao, Rui-Lin; Lumyong, Saisamorn; Boonmee, Saranyaphat; Wen, Ting-Chi; Mortimer, Peter E.; Xu, Jianchu — Fungal diversity notes 929–1035: taxonomic and phylogenetic contributions on genera and species of fungi — Cytospora ulmicola

Uraian asli dan riwayat sumber

Riwayat atau halaman sumber

Ringkasan sumber

Associated with twigs and branches of Ulmus pumila L. (Ulmaceae). Sexual morph Undetermined. Asexual morph Conidiostromata 1000–1700 × 500–800 µm diam., semi-immersed in host tissue, solitary, erumpent, scattered, discoid, circular to ovoid, with multi-loculate, pycnidial, embedded in stromatic tissue, with ostiole. Ostioles 200–500 µm long, with an ostiolar neck. Conidiomata walls comprising a few layers of pseudoparenchymatous cells of textura angularis, with inner layer thin, pale brown, outer layer brown to dark brown.

Revisi sumber

Morfologi sumber

Associated with twigs and branches of Ulmus pumila L. (Ulmaceae). Sexual morph Undetermined. Asexual morph Conidiostromata 1000–1700 × 500–800 µm diam., semi-immersed in host tissue, solitary, erumpent, scattered, discoid, circular to ovoid, with multi-loculate, pycnidial, embedded in stromatic tissue, with ostiole. Ostioles 200–500 µm long, with an ostiolar neck. Conidiomata walls comprising a few layers of pseudoparenchymatous cells of textura angularis, with inner layer thin, pale brown, outer layer brown to dark brown. Conidiophores unbranched or occasionally branched at the base, formed from the inner layer of the pycnidial wall. Conidiogenous cells (6–)9–14 × 1.3–1.8(–2.8) μm (\(\bar{x} \) = 11 × 2 μm, n = 15), blastic, enteroblastic, flask-shaped, phialidic, hyaline, and smooth-walled. Conidia (7–)8–9.5 × 1.5–1.7(–1.9) µm (\(\bar{x} \) = 8.5 × 1.6 µm, n = 30), unicellular, hyaline, oblong to allantoid, smooth-walled. Culture characteristics: Conidia germinating on PDA within 12 h. Germ tubes produced from all sides. Colonies on PDA reaching 22–27 mm diam. after 2 weeks at room temperature, colonies irregular in shape, medium dense, flat to slightly raised, surface slightly rough, with small granular, to velvety, edge undulate with well-defined margin, from above pale orange to light brown at the margin, amber near the margin, with, zonate, pale yellowish to dandelion and dark amber at the centre; from below pale yellowish at the margin, with dark amber at the centre; not producing pigmentation in agar.

Revisi sumber

Ekologi sumber

Culture characteristics: Conidia germinating on PDA within 12 h. Germ tubes produced from all sides. Colonies on PDA reaching 22–27 mm diam. after 2 weeks at room temperature, colonies irregular in shape, medium dense, flat to slightly raised, surface slightly rough, with small granular, to velvety, edge undulate with well-defined margin, from above pale orange to light brown at the margin, amber near the margin, with, zonate, pale yellowish to dandelion and dark amber at the centre; from below pale yellowish at the margin, with dark amber at the centre; not producing pigmentation in agar. Material examined: RUSSIA, Rostov region, Shakhty City, street trees, on dead branch of Ulmus pumila L. (Ulmaceae), 27 May 2017, T.S. Bulgakov, T-1778 (MFLU 17-2080, holotype; PDD, isotype), ex-type living culture, MFLUCC 18-1227. Notes: Cytospora ulmicola (MFLU 17-2080) was found on a dead branch (with signs of necrosis) of Ulmus pumila in European Russia. Cytospora ulmicola resembles Cytospora species in having an ostiolar neck with multi-loculate conidiostromata and unicellular, oblong to allantoid conidia. Phylogenetic analysis based on a combined ITS, LSU and ACT sequence dataset indicates that C. ulmicola clusters with C. cotini Norph. et al. (MFLUCC 14-1050), C. ampulliformis Norph. et al. (MFLUCC 16-0583), C. gelida Norph. et al. (MFLUCC 16-0634), and C. ceratosperma (Tode) G.C.

Revisi sumber

Struktur reproduksi sumber

Associated with twigs and branches of Ulmus pumila L. (Ulmaceae). Sexual morph Undetermined. Asexual morph Conidiostromata 1000–1700 × 500–800 µm diam., semi-immersed in host tissue, solitary, erumpent, scattered, discoid, circular to ovoid, with multi-loculate, pycnidial, embedded in stromatic tissue, with ostiole. Ostioles 200–500 µm long, with an ostiolar neck. Conidiomata walls comprising a few layers of pseudoparenchymatous cells of textura angularis, with inner layer thin, pale brown, outer layer brown to dark brown. Conidiophores unbranched or occasionally branched at the base, formed from the inner layer of the pycnidial wall. Conidiogenous cells (6–)9–14 × 1.3–1.8(–2.8) μm (\(\bar{x} \) = 11 × 2 μm, n = 15), blastic, enteroblastic, flask-shaped, phialidic, hyaline, and smooth-walled.

Revisi sumber

Rujukan halaman ini

Fungi. Cytospora ulmicola. Rilis data fungi-2026-10-12-expansion-2. https://fungi-atlas.mgadib.chatgpt.site/jamur/cytospora-ulmicola/.

Status: impor sumber; pemeriksaan editorial per profil belum selesai. Untuk klaim biologis, gunakan rujukan pada bagian terkait dan periksa konteks sumber aslinya.